The way I understand it: It was designed to flow a certain amount. In other words it can handle a certain amount of water (volume and flow rate) forced through before they feel serious damage will occur. However, there is nothing stopping more water from being forced through it than the original designers thought might happen. This can occur because the head between the two rivers has increased over time. Just like there is nothing stopping the guy running the mud pumps from overpressuring the pipe. Just because it's rated for 10,000 psi or whatever doesn't mean I can't force more through it and break it. In this case God is running the pumps. Maybe a bad analogy but the best I could come up with quickly.
In our case though, the 620,000 cfs orignal design capacity has been increased by adding two more structures so the actual capacity is more like 1.14 million. It appears they have left the 620,000 cfs design limit on the structurs and use the new added capacity as safety factor.